Aquatic and Wetland Vascular Plants of the Northern Great Plains
4. Polypodiaceae, the Polypody Family
3. Dryopteris Adans. -- Shield fern1. Dryopteris carthusiana (Vill.) H. P. Fuchs -- Spinulose woodfern
Fronds alike, clustered, to 10 dm tall, the blades bipinnate-pinnatifid to nearly tripinnate, lanceolate, 2-6 dm long, 6-30 cm wide, acuminate, slightly narrowed at the base; pinnae usually numbering 10-15 pairs, ascending to spreading, alternate to subopposite, lanceolate to linear-lanceolate; pinnules toothed to deeply pinnately lobed, mostly 5-40 mm long, 3-10 mm wide, the teeth or lobes spinulose-tipped. Jun--Sep. Wet alluvial woods or swamps; uncommon; e ND, n and se NE; (Labr. to Alta. s to VA, MO, NE and ID). D. austriaca (Jacq.) Woynar var. spinulosa (Mull.) Fiori; D. spinulosa (Mull.) Watt.
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