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Winter Severity and Wolf Predation on
a Formerly Wolf-free Elk Herd


Our study was supported by National Geographic Society, Yellowstone National Park, and the Biological Resources Division of the U.S. Geological Survey. We thank the following volunteers and technicians who assisted with the study: M. E. Nelson, G. D. DelGiudice, C. A. Campbell, D. Guernsey, D. R. Stahler, S. Evans, T. Zeiber, and K. Honness; R. Stradly who safely piloted the aircraft; W. Brewster and J. Varley, Yellowstone National Park, for administrative oversight; and L. Diekmann, Yellowstone Park Foundation, for fiscal management. We also appreciate the suggestions made by D. Maehr and the anonymous reviewers who greatly improved the paper.
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